We thank Beth Sanderson and Mindi Sheer, NOAA’s NW Fisheries Science Center, and Lucinda
Johnson, University of Minnesota, for careful and constructive manuscript reviews that substantially
improved the manuscript. Sean Luis provided endless hours of assistance tracking down and formatting
references. This manuscript was stimulated by presentations during, and discussions following, the 2008
annual meeting of the International Association of Landscape Ecology in Wageningen, The Netherlands.
Partial funding for manuscript preparation was provided by U.S. Environmental Protection Agency grant
RM832827 to Oregon State University, the Austrian – American Education Commission, and the Fulbright